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Psychedelics paired with therapy could treat chronic mental health conditions Research and Innovation Certainly these illusory effects could arise through a variety of mechanisms, because 5-HT2A receptors are expressed in many areas of the brain responsible for cognition and sensory processing. It has not been widely appreciated, however, that mammalian primary visual cortex area V1, the largest known visual cortical area, expresses a high density of 5-HT2A receptors (Watakabe et al., 2009; Moreau et al., 2010). Microiontophoresis of DOI into macaque V1 gave a bidirectional modulatory effect on the neuron’s firing rate. Analysis of recordings from 44 neurons showed that DOI facilitates visual responses of neurons with a low firing rate but suppressed those of neurons with a high firing rate. The authors suggest that neurons in the input layers of V1, which abundantly express the 5-HT2A receptor, may act as gain controllers by enhancing weak signal response and suppressing excessive r